Hyaenodon
horridus
Click
here to see the skeleton of Hyaenodon
The Creodonts (flesh-tooth) are a long
extinct group of carnivorous mammals whose rise as meat eaters is an example of parallel
evolution to modern carnivores; the Fissipedia.
Creodonts are thought to have
evolved from primitive insectivores of the late Cretaceous. The variety of sizes
and forms that Creodonts achieved rival that of the Fissipeds. Sometime during
the Paleocene, the Creodonts branched into two families; the Oxyaenidae, which
were rather catlike in form, and the Hyaenodontidae, which were more dog like. The
order became extinct in the Pliocene epoch, perhaps unable
to compete with the rapidly evolving felids, canids and ursids. Not to mention the increasing
adaptation towards speed in prey species, most notably in the Artiodactyls (
Even-toed herbivores).
Creodonts are characterized
by relatively unspecialized features in comparison to today's carniores. The teeth, which are obviously adapted for
flesh eating, are rather morphologically conservative when comparing them to
modern Fissipeds. They did have carnassial ( flesh-shearing ) molars, but these were placed further back in the tooth row
than in modern carnivora. Click here to see the skeleton
Other features of
many Creodonts
were the short stocky limbs, and the five toed feet (pentadactylus) which ended
in short, blunt, hoof-like claws. Though not terribly speedy creodonts were
probably very active hunters.
Hyaenodon
horridus, is considered to be a cursorial type, and therefore was an effective
predator. But still unable to survive past the Pliocene.
Hyaenodon
horridus is a
common predator from the Oligocene epoch of western North America. Its remains
are found in the Chadron and Brule formations of the Nebraska and South Dakota
badlands. Hyaenodon was wolf sized. It had a long, low lumbar ( back ) region, a
primitive feature that it shared with many Creodonts and other groups of
ancestral early mammals.
Hyaenodons skull was long and
massive and filled with large bone-crushing teeth. But its brain case was very
small, so the huge skull must have served mostly to anchor large jaw operating
muscles. Like all Creodonts, the skull had no ossified ( bony ) auditory bulla. These
bulbous structures are found on the rear underside of the skull in many modern mammals, and housed the delicate bones
of the inner ear. This structure was probably cartilaginous in Creodonts and
thus was not fossilized.
This painting depicts two
Hyaenodon approaching the carcass of a Merycoidodon. An Oreodont whose numerous
fossil remains tell us that these herbivores must have roamed the pre-badlands
plains in large numbers.
The scavenging bird shown
here is a hypothetical crow-like opportunist.
Because of the fragile nature
of their hollow bones, bird fossils are very rare in
any time period of the geologic record since their appearance in the Jurassic.
Avian species that are known to exist during the Oligocene world include;
rails, Grebes, flamingos, woodpeckers, hawks, eagles, vultures, limpkins,
and cranes. This
evidence shows that many modern forms were already well established at this time.
Though this record so far does not include crows or other smaller scavenger
types, there surely
must have been species to fill this niche during the Oligocene. This
is how I justify the depiction of an animal whose existence is not yet supported by
fossil evidence.
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